As a result, this game lacks much of the sense of humor of its predecessors.
Morden's infantry, Morden himself and Allen O'Neil are nowhere to be found in this game rather, one fights armies of veiled terrorists, special forces, and assorted mecha. 'Fat mode' is the only remaining modifier, there are no zombies, mummies, aliens or other supernatural creatures, although the final boss is a large demon. The player may now perform a slide-dash. At the end of the game, the Ptolemaic Army summons a giant demon as the final boss, which after a long battle is forced to leave Earth thanks to the heroes. Together once more, the quartet investigate the shrouded objective of the Ptolemaic Army, who over time grows more powerful as they are joined by a mysterious masked man and his followers. Marco and Tarma of the Peregrine Falcon Strike Force follow in hot pursuit against the group and in the process are joined by Eri and Fio of SPARROWS. 
A special disc that contains deep and intricate secrets about the Metal Slug project is stolen by a mysterious group called the Ptolemaic Army, whose speciality lies from within archaeological excavation and espionage. One year after the event of Metal Slug 7.
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